|  | Utricularia graminifolia – the carnivorous 
        foreground plant |   |  | 
     
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      |  | Article by Ole Pedersen, Troels Andersen and 
        Claus Christensen |  |  |  | 
     
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      |  | Utricularia 
        graminifolia (Tropica No 049B) belongs to the most interesting water plants. 
        It is carnivorous and creates an attractive green carpet in the foreground 
        within a couple of months. Utricularia graminifolia may be difficult to 
        start up but once it has gotten a firm grip in the aquarium, it offers 
        a unique experience for the aquarist. The tiny trapping devices may inspire 
        long discussions among family members and friends as carnivorous plants 
        always stimulate people’s curiosity. |  | 
     
      |  |  | Utricularia 
        graminifolia in its natural habitat on the edge of a shaded stream in 
        southern Vietnam. The plant grows partly submerged and partly emerged 
        among the leaf litter. |  | 
     
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      |  | Utricularia 
        graminifolia belongs to the bladderwort family (Lentibulariaceae) and 
        originates in SE Asia where it has been found in the southern China, India, 
        Sri Lanka, Burma, Laos and Vietnam. At those sites, it appears as amphibious 
        swamp plant as well as totally submerged in springs and along small streams. 
        Its natural habitat is often shaded and it rarely grows in full sunlight. 
        It always grew in very soft water where we found it and we had severe 
        difficulties keeping it alive during transportation. Fortunately, the 
        plant that is being produced now is much sturdier and thus, more suitable 
        for use in aquaria but it remains a plant for the specialist. Utricularia 
        aurea is another bladderwort which occasionally is found in the shops 
        but it looks completely different with a growth form similar to species 
        of Ceratophyllum and Myriophyllum. |  | 
     
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      |  |  | Close-up of 
        the trapping devices on Utricularia graminifolia that are used to catch 
        crustaceans and ciliates. The animals contain nitrogen and phosphorous 
        that can be used by the plant after the tissue has been dissolved by enzymes 
        in the tiny traps. Photo by Oliver Knott. |  | 
     
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      |  | Bladderworts 
        are all carnivorous plants and many of them are associated with water. 
        The genus name “Utricularia” refers to the bladders (trapping devices) 
        whereas the species name “graminifolia” refers to the grass-leaved leaves. 
        The grass-leaved leaves are exactly what distinguish it from the other 
        approximately 210 species of bladderworts that have been described so 
        far. The tiny grass-leaved leaves form an imbricate structure of freshly 
        green leaves, which normally covers the bladders that are formed on the 
        rhizome and the midribs. It flowers when it grows emerged. |  | 
     
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      |  | However, what 
        makes Utricularia graminifolia unique to the aquarist is the tiny 2 mm 
        long bladders that in nature traps crustaceans and ciliates seeking shelter 
        among leaves and rhizomes. The animals are digested by enzymes secreted 
        by the into the bladders, and the released nutrients are subsequently 
        taken up by the plant and used to support new growth. Utricularia graminifolia 
        often grows in very nutrient poor environments and thus, it is a huge 
        advantage to be able to explore alternative nutrient sources of particularly 
        nitrogen (N) and phosphorous (P) by trapping nutritious animals. |  | 
     
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      |  |  | Here, Utricularia 
        graminifolia is used in aquascaping where it forms a green stream in the 
        aquarium. The plant may also be used as ordinary foreground plant where 
        it forms a continuous green carpet 6-8 week after planting. Photo by Oliver 
        Knott. |  | 
     
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      |  | Tropica 
        grows Utricularia graminifolia emergent and it is supplied on either stone 
        wool or coir discs. When planted in the aquarium, the clod should be divided 
        into 6 to 8 smaller pieces and planted with a distance of 5 cm in the 
        foreground of the aquarium or on a slope in the aquascape. It is important 
        to keep about 1 cm of the coir or stone wool in order to provide good 
        anchoring of the plants. Using tweezers to insert the tiny blocks of plants 
        makes the entire process of planting much easier. To begin with, Utricularia 
        graminifolia should be offered relatively low light but after the initial 
        6-8 weeks, where after the plants have obtained a firm hold in the aquarium, 
        the light may be increased. At this point in time, the tiny blocks have 
        now been transformed into a continuous grass-like carpet and this is the 
        time where PLANT NUTRITION liquid should be supplied in order to prevent 
        iron and manganese deficiencies. Our experience with Utricularia graminifolia 
        shows that it is doing best at a pH of 6.8 to 7.0 in water of 7-10 dKh 
        hardness. |  | 
     
      |  | Fertilization 
        with CO2 is not required but if supplied, the growth is stimulated significantly. 
        Although the commercially available variety is sturdier that the wild 
        type, the initial phase after planting may prove critical since the Utricularia 
        graminifolia is sensitive to environmental changes. Thus, make sure that 
        the recommendations are followed and keep other fast-growing plants out 
        of the new patches with Utricularia graminifolia until the plants are 
        well established in the aquarium. Utricularia graminifolia is relatively 
        fragile and thus, it should not be kept with fishes with burrowing behavior. |  | 
     
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